Galeopterus variegatus

(G. Cuvier, 1797)

Sunda Colugo

Taxonomy

Subclass : Theria
Infraclass : Placentalia
Magnorder : Boreoeutheria
Superorder : Euarchontoglires
Order : Dermoptera
Family : Cynocephalidae
Genus : Galeopterus

Species status

Living
Found in the wild
Listed in MSW3 2005

Authority citation

Cuvier, G. 1797-12-24. Tableau élémentaire de l'histoire naturelle des animaux. Baudouin, Paris, 710 pp.

Authority publication link

https://www.biodiversitylibrary.org/page/11637189

Original name as described

Galeopithecus variegatus

Other common names

Sunda Flying Lemur · Bornean Colugo · Javan Colugo · Malayan Colugo · Sumatran Colugo

Type locality

type locality not given. Restricted by O. Thomas in 1908 to the "Malay Peninsula and Islands."

Biogeographic realm

Indomalaya

Country distribution

Laos · Vietnam · Cambodia · Myanmar · Thailand · Malaysia · Singapore · Indonesia

Taxonomy notes

there appears to be strong molecular and morphological evidence suggesting there are at least 6 diagnosable species currently included under G. variegatus, although no taxonomic changes have been made yet; further research is needed to determine the scientific names that apply to each of the 6 putative species

Taxonomy notes citation

Janečka, J. E., Helgen, K. M., Lim, N. T. L., Baba, M., Izawa, M., & Murphy, W. J. (2008). Evidence for multiple species of Sunda colugo. Current Biology, 18(21), R1001-R1002. · Mason, V. C., Li, G., Minx, P., Schmitz, J., Churakov, G., Doronina, L., ... & Murphy, W. J. (2016). Genomic analysis reveals hidden biodiversity within colugos, the sister group to primates. Science advances, 2(8), e1600633. · Mason, V. C. (2016). Reassessing Colugo Phylogeny, Taxonomy, and Biogeography by Genome Wide Comparisons and DNA Capture Hybridization from Museum Specimens (Doctoral dissertation).

IUCN Red List status

Least Concern

Species Permalink

https://www.mammaldiversity.org/taxon/1000535

Country distribution map

The map below provides a general overview. Most species inhabit only specific regions within countries. Hover over the highlighted regions to see the country name.

Names and synonyms

Present and past (if available) associated names to the species. Click on a name to view its details. If the list is long, scroll down to see more.

Galeopithecus Temminckii

(G. R. Waterhouse, 1839)

Family

Cynocephalidae

Root name

temminckii

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1907.1.1.220

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/259a10df-1df7-4cc3-88d2-90747b54c796

Authority page

119

Authority page URI

https://www.biodiversitylibrary.org/page/30571180

Authority publication

Proceedings of the Zoological Society of London

Name usages

  1. Lesson (1842:11) (information at https://hesperomys.com/a/36812)
  2. Smith (1842:112) (information at https://hesperomys.com/a/67141)
  3. Stafford (2005) (information at https://hesperomys.com/a/9708)

Cynocephalus tellonis

(Lyon, 1908)

Family

Cynocephalidae

Root name

tellonis

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1907.6.18.2

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/56992928-d5f8-442d-8e5c-8bd6fb53122d

Authority page

139

Authority page URI

https://www.biodiversitylibrary.org/page/24295848

Authority publication

Annals and Magazine of Natural History

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus taylori

O. Thomas, 1908

Family

Cynocephalidae

Root name

taylori

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1908.1.25.3

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/1c347c2b-aade-476f-b1f3-6fc9fea6fb50

Authority page

102

Authority page URI

https://www.biodiversitylibrary.org/page/19042812

Authority publication

Journal of the Federated Malay States Museums

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus peninsulae

O. Thomas, 1908

Family

Cynocephalidae

Root name

peninsulae

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1908.7.20.10

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/546466ff-19c4-4369-af17-94a5b0fceb4d

Authority page

303

Authority page URI

https://www.biodiversitylibrary.org/page/22071072

Authority publication

Annals and Magazine of Natural History

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus variegatus terutaus

Chasen & Kloss, 1929

Family

Cynocephalidae

Root name

terutaus

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1947.1427 (= FMSM 264/17)

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/93b3e643-0862-44eb-abfc-3e0b6305ffcc

Authority page

11

Authority publication

Bulletin of Raffles Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus variegatus perhentianus

Chasen & Kloss, 1929

Family

Cynocephalidae

Root name

perhentianus

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1947.1428

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/a2e2a357-caef-4e88-a4f3-1c81457523dd

Authority page

11

Authority publication

Bulletin of Raffles Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus hantu

Cabrera, 1924

Family

Cynocephalidae

Root name

hantu

Validity status

synonym

Nomenclatural status

available

Type

MNCN 20-II-19-9

Type kind

holotype

Authority page

128

Authority publication

Boletín de la Real Sociedad Española de Historia Natural

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus rufus

(G. Cuvier, 1797)

Family

Cynocephalidae

Root name

rufus

Validity status

synonym

Nomenclatural status

available

Type

MNHN 224

Type kind

holotype

Authority page

107

Authority page URI

https://www.biodiversitylibrary.org/page/11637189

Authority publication

Paris

Name usages

  1. Cuvier (1797:107, https://www.biodiversitylibrary.org/page/11637189) (information at https://hesperomys.com/a/38531)
  2. Fischer (1814:xviii, https://www.biodiversitylibrary.org/page/29130079) (information at https://hesperomys.com/a/12226)
  3. Lesson (1827:75, https://www.biodiversitylibrary.org/page/54207543) (information at https://hesperomys.com/a/36839)
  4. Lesson (1842:11) (information at https://hesperomys.com/a/36812)
  5. Fitzinger (1855:120) (information at https://hesperomys.com/a/65926)

Galeopterus lechei

Gyldenstolpe, 1920

Family

Cynocephalidae

Root name

lechei

Validity status

synonym

Nomenclatural status

available

Type

NHMO 7986 (= NHMO 24)

Type kind

holotype

Authority page

17

Authority page URI

https://www.biodiversitylibrary.org/page/41920242

Authority publication

Kunglika Svenska Vetenskapsakademiens Handlingar

Galeopithecus pumilus

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

pumilus

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:104448

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/33f0039e0-3bd9-4b89-b6bb-c92225ee2e62

Authority page

46

Authority page URI

https://www.biodiversitylibrary.org/page/8809960

Authority publication

Smithsonian Collections

Galeopithecus gracilis

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

gracilis

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:104601

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/37649af06-f8f5-4dba-a372-12a39faa26c0

Authority page

49

Authority page URI

https://www.biodiversitylibrary.org/page/8809967

Authority publication

Smithsonian Collections

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus natunae

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

natunae

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:104602

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/3fed9202d-0421-47aa-8f1b-5ad505bb9b7f

Authority page

50

Authority page URI

https://www.biodiversitylibrary.org/page/8809968

Authority publication

Smithsonian Collections

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus aoris

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

aoris

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:112428

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/35e78b160-3463-406e-8933-1c3b21783103

Authority page

47

Authority page URI

https://www.biodiversitylibrary.org/page/8809963

Authority publication

Smithsonian Collections

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus tuancus

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

tuancus

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:114375

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/3573347df-82ce-4833-b297-5d7f30fe378f

Authority page

53

Authority page URI

https://www.biodiversitylibrary.org/page/8809975

Authority publication

Smithsonian Collections

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus saturatus

(G. S. Miller, 1903)

Family

Cynocephalidae

Root name

saturatus

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:121750

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/3dcf8dbac-8c63-4e31-af54-d9ea349c6638

Authority page

51

Authority page URI

https://www.biodiversitylibrary.org/page/8809971

Authority publication

Smithsonian Collections

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus chombolis

Lyon, 1909

Family

Cynocephalidae

Root name

chombolis

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:144375

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/36222ee63-2094-481b-8702-465b136b8341

Authority page

486

Authority page URI

https://www.biodiversitylibrary.org/page/15714615

Authority publication

Proceedings of the United States National Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus abbotti

Lyon, 1911

Family

Cynocephalidae

Root name

abbotti

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:145577

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/3bff21b75-039a-4c9f-bba6-00839a7904b4

Authority page

126

Authority page URI

https://www.biodiversitylibrary.org/page/15845288

Authority publication

Proceedings of the United States National Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus lautensis

Lyon, 1911

Family

Cynocephalidae

Root name

lautensis

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:151886

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/304435280-a25f-4804-891d-2343820fbef9

Authority page

125

Authority page URI

https://www.biodiversitylibrary.org/page/15845287

Authority publication

Proceedings of the United States National Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus borneanus

Lyon, 1911

Family

Cynocephalidae

Root name

borneanus

Validity status

synonym

Nomenclatural status

available

Type

USNM:MAMM:151888

Type kind

holotype

Type specimen URI

http://n2t.net/ark:/65665/383ba50ef-7ea3-4193-8972-cf96921ebb34

Authority page

124

Authority page URI

https://www.biodiversitylibrary.org/page/15845286

Authority publication

Proceedings of the United States National Museum

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Cynocephalus variegatus

(Corbet & J. Edwards Hill, 1980)

Family

Cynocephalidae

Root name

variegatus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

35

Authority publication

London

Name usages

  1. Corbet & Hill (1980:35) (information at https://hesperomys.com/a/63069)
  2. Honacki, Kinman & Koeppl (1982:110) (information at https://hesperomys.com/a/63071)

lecheyi

Stafford, 2005

Family

Cynocephalidae

Root name

lecheyi

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

110

Authority publication

Baltimore

Name usages

Stafford (2005:110) (information at https://hesperomys.com/a/9708)

pumilis

Stafford, 2005

Family

Cynocephalidae

Root name

pumilis

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

110

Authority publication

Baltimore

Name usages

Stafford (2005:110) (information at https://hesperomys.com/a/9708)

Galeopterus variegatus

Stafford, 2005

Family

Cynocephalidae

Root name

variegatus

Validity status

synonym

Nomenclatural status

name_combination

Authority publication

Baltimore

Name usages

  1. Stafford (2005) (information at https://hesperomys.com/a/9708)
  2. Mammal Diversity Database (2018:ID #41502) (information at https://hesperomys.com/a/67336)
  3. Wilson & Mittermeier (2018:285) (information at https://hesperomys.com/a/57916)
  4. Mammal Diversity Database (2019:ID #41502) (information at https://hesperomys.com/a/67337)
  5. Mammal Diversity Database (2024, https://www.mammaldiversity.org/taxon/1000535) (information at https://hesperomys.com/a/67250)
  6. International Union for the Conservation of Nature (2024, https://www.iucnredlist.org/species/41502/10479343) (information at https://hesperomys.com/a/67246)

Galeopterus variegatus borneanus

D. E. Wilson & Mittermeier, 2018

Family

Cynocephalidae

Root name

borneanus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

285

Authority publication

Barcelona

Name usages

Wilson & Mittermeier (2018:285) (information at https://hesperomys.com/a/57916)

Galeopterus variegatus peninsulae

D. E. Wilson & Mittermeier, 2018

Family

Cynocephalidae

Root name

peninsulae

Validity status

synonym

Nomenclatural status

name_combination

Authority page

285

Authority publication

Barcelona

Name usages

Wilson & Mittermeier (2018:285) (information at https://hesperomys.com/a/57916)

Galeopterus variegatus temminckii

D. E. Wilson & Mittermeier, 2018

Family

Cynocephalidae

Root name

temminckii

Validity status

synonym

Nomenclatural status

name_combination

Authority page

285

Authority publication

Barcelona

Name usages

Wilson & Mittermeier (2018:285) (information at https://hesperomys.com/a/57916)

Galeopterus variegatus variegatus

D. E. Wilson & Mittermeier, 2018

Family

Cynocephalidae

Root name

variegatus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

285

Authority publication

Barcelona

Name usages

Wilson & Mittermeier (2018:285) (information at https://hesperomys.com/a/57916)

Galeopithecus variegatus

(G. Cuvier, 1797)

Family

Cynocephalidae

Root name

variegatus

Validity status

species

Nomenclatural status

available

Authority page

107

Authority page URI

https://www.biodiversitylibrary.org/page/11637189

Authority publication

Paris

Name usages

  1. Cuvier (1797:107, https://www.biodiversitylibrary.org/page/11637189) (information at https://hesperomys.com/a/38531)
  2. Fischer (1814:xviii, https://www.biodiversitylibrary.org/page/29130079) (information at https://hesperomys.com/a/12226)
  3. Lesson (1827:75, https://www.biodiversitylibrary.org/page/54207543) (information at https://hesperomys.com/a/36839)

Galeopithecus rufus

(J. B. Audebert, 1799)

Family

Cynocephalidae

Root name

rufus

Validity status

synonym

Nomenclatural status

preoccupied

Authority page

35

Authority page URI

https://www.biodiversitylibrary.org/page/58948949

Authority publication

Paris

Galeopithecus variegatus

(J. B. Audebert, 1799)

Family

Cynocephalidae

Root name

variegatus

Validity status

synonym

Nomenclatural status

subsequent_usage

Authority page

37

Authority page URI

https://www.biodiversitylibrary.org/page/58948953

Authority publication

Paris

Galeopithecus ternatensis

(A. G. Desmarest, 1817)

Family

Cynocephalidae

Root name

ternatensis

Validity status

synonym

Nomenclatural status

available

Authority page

377

Authority page URI

https://www.biodiversitylibrary.org/page/18058139

Authority publication

Nouveau Dictionnaire d'Histoire Naturelle

Name usages

  1. Lesson (1827:75, https://www.biodiversitylibrary.org/page/54207543) (information at https://hesperomys.com/a/36839)
  2. Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus varius

(A. G. Desmarest, 1817)

Family

Cynocephalidae

Root name

varius

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

376

Authority page URI

https://www.biodiversitylibrary.org/page/18058136

Authority publication

Nouveau Dictionnaire d'Histoire Naturelle

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus rufus

(A. G. Desmarest, 1821)

Family

Cynocephalidae

Root name

rufus

Validity status

synonym

Nomenclatural status

subsequent_usage

Authority page

108

Authority page URI

https://www.biodiversitylibrary.org/page/39523872

Authority publication

Paris

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopithecus marmoratus

(Temminck, 1828)

Family

Cynocephalidae

Root name

marmoratus

Validity status

synonym

Nomenclatural status

nomen_nudum

Authority page

66 (footnote)

Authority publication

Bijdragen aan de Natuurkundige Wetenschappen

Galeopithecus marmoratus

(Temminck, 1836)

Family

Cynocephalidae

Root name

marmoratus

Validity status

synonym

Nomenclatural status

nomen_nudum

Authority page

9

Authority page URI

https://www.biodiversitylibrary.org/page/51389725

Authority publication

Leiden

Galeopithecus undatus

(J. A. Wagner, 1840)

Family

Cynocephalidae

Root name

undatus

Validity status

synonym

Nomenclatural status

available

Authority page

326

Authority page URI

https://www.biodiversitylibrary.org/page/31071965

Authority publication

Die Säugthiere in Abbildungen nach der Natur

Name usages

Stafford (2005) (information at https://hesperomys.com/a/9708)

Galeopterus lautenis

Chasen, 1940

Family

Cynocephalidae

Root name

lautenis

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

21

Authority publication

Bulletin of Raffles Museum

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