Callosciurus pygerythrus

(I. Geoffroy Saint-Hilaire, 1831)

Irrawaddy Squirrel

Taxonomy

Subclass : Theria
Infraclass : Placentalia
Magnorder : Boreoeutheria
Superorder : Euarchontoglires
Order : Rodentia
Suborder : Sciuromorpha
Family : Sciuridae
Subfamily : Callosciurinae
Tribe : Callosciurini
Genus : Callosciurus

Species status

Living
Found in the wild
Listed in MSW3 2005

Authority citation

Geoffroy Saint-Hilaire, I. 1831-03-19. Mammifères. Pp. 1–160 in Bélanger, C. 1835. Voyage aux Indes-Orientales, par le Nord de l'Europe, les Provinces du Caucase, la Géorgie, l'Arménie et la Perse, suivi de Détails topographiques, statistiques et autres sur le Pégou, les Iles de Java, de Maurice et de Bourbon, sur le Cap de Bonne-Espérance et Sainte-Hélène, pendant les Années 1825, 1826, 1827, 1828 et 1829. Zoologie. Arthus Bertrand, Paris, 535 pp.

Authority publication link

https://hdl.handle.net/2027/osu.32435028294999

Original name as described

Sciurus pygerythrus

Other common names

Hoary-bellied Squirrel

Type material

MNHN 1829-286 (= MNHN 294)

Type kind

holotype

Type locality

"dans les forêts de Syriam, au Pégou," Myanmar.

Biogeographic realm

Palearctic · Indomalaya

Country distribution

Nepal · India · Bhutan · Bangladesh · Myanmar · China

IUCN Red List status

Least Concern

Species Permalink

https://www.mammaldiversity.org/taxon/1001548

Country distribution map

The map below provides a general overview. Most species inhabit only specific regions within countries. Hover over the highlighted regions to see the country name.

Names and synonyms

Present and past (if available) associated names to the species. Click on a name to view its details. If the list is long, scroll down to see more.

Sciurus pygerythrus

(I. Geoffroy Saint-Hilaire, 1831)

Family

Sciuridae

Root name

pygerythrus

Validity status

species

Nomenclatural status

available

Type

MNHN 1829-286 (= MNHN 294)

Type kind

holotype

Authority page

145

Authority publication

Paris

Name usages

  1. Geoffroy Saint-Hilaire (1831:145) (information at https://hesperomys.com/a/67807)
  2. Blyth (1849:602, https://www.biodiversitylibrary.org/page/40128631) (information at https://hesperomys.com/a/63252)
  3. Blyth (1862:334 (footnote), https://www.biodiversitylibrary.org/page/37181614) (information at https://hesperomys.com/a/68511)
  4. Blyth (1863:103, https://www.biodiversitylibrary.org/page/47534784) (information at https://hesperomys.com/a/37155)
  5. Blyth (1875:37, https://www.biodiversitylibrary.org/page/38675003) (information at https://hesperomys.com/a/68967)
  6. Thomas (1908:246, https://www.biodiversitylibrary.org/page/2222283) (information at https://hesperomys.com/a/62341)

Sciurus Lokroides

(B. H. Hodgson, 1836)

Family

Sciuridae

Root name

lokroides

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1843.1.12.58

Type kind

lectotype

Type specimen URI

https://data.nhm.ac.uk/object/3813ce5c-8ba9-4dd5-8531-7399cca54b8e

Authority page

232

Authority page URI

https://www.biodiversitylibrary.org/page/37188329

Authority publication

Journal of the Asiatic Society of Bengal

Name usages

  1. Hodgson (1836:232, https://www.biodiversitylibrary.org/page/37188329) (information at https://hesperomys.com/a/68654)
  2. Ogilby (1839:lxvii, https://www.biodiversitylibrary.org/page/2912378) (information at https://hesperomys.com/a/67483)
  3. Blyth (1847:873, https://www.biodiversitylibrary.org/page/40126422) (information at https://hesperomys.com/a/39852)
  4. Horsfield (1857:402, https://www.biodiversitylibrary.org/page/12860976) (information at https://hesperomys.com/a/36026)
  5. Blyth (1863:104, https://www.biodiversitylibrary.org/page/47534785) (information at https://hesperomys.com/a/37155)
  6. Murray (1866:353, https://www.biodiversitylibrary.org/page/15580349) (information at https://hesperomys.com/a/39798)
  7. Jerdon (1867:169, https://www.biodiversitylibrary.org/page/37299942) (information at https://hesperomys.com/a/68353)

Funambulus lokrioïdes

(Lesson, 1842)

Family

Sciuridae

Root name

lokrioides

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

110

Authority publication

Paris

Name usages

Lesson (1842:110) (information at https://hesperomys.com/a/36812)

Funambulus Pygerythrus

(Lesson, 1842)

Family

Sciuridae

Root name

pygerythrus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

109

Authority publication

Paris

Name usages

Lesson (1842:109) (information at https://hesperomys.com/a/36812)

Sciurus Assamensis

(J. E. Gray, 1843)

Family

Sciuridae

Root name

assamensis

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1879.11.21.584

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/938bcc82-1922-4b3c-921f-b310873fde00 https://data.nhm.ac.uk/object/938bcc82-1922-4b3c-921f-b310873fde00/1686614400000

Authority page

143

Authority page URI

https://www.biodiversitylibrary.org/page/53729844

Authority publication

British Museum Catalogue

Name usages

  1. Blyth (1862:334 (footnote), https://www.biodiversitylibrary.org/page/37181614) (information at https://hesperomys.com/a/68511)
  2. Blyth (1863:103, https://www.biodiversitylibrary.org/page/47534784) (information at https://hesperomys.com/a/37155)
  3. Murray (1866:353, https://www.biodiversitylibrary.org/page/15580349) (information at https://hesperomys.com/a/39798)
  4. Blyth (1875:37, https://www.biodiversitylibrary.org/page/38675003) (information at https://hesperomys.com/a/68967)
  5. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)

Sciurus Locroides

(B. H. Hodgson, 1844)

Family

Sciuridae

Root name

locroides

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

293

Authority page URI

https://www.biodiversitylibrary.org/page/46405528

Authority publication

Calcutta Journal of Natural History

Name usages

Hodgson (1844:293, https://www.biodiversitylibrary.org/page/46405528) (information at https://hesperomys.com/a/65716)

Sciurus blythii

(Tytler, 1854)

Family

Sciuridae

Root name

blythii

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1879.11.21.361

Type kind

lectotype

Type specimen URI

https://data.nhm.ac.uk/object/deb5f5f7-8f8c-4cb5-9d13-d79d330aad31

Authority page

172

Authority page URI

https://www.biodiversitylibrary.org/page/18659631

Authority publication

Annals and Magazine of Natural History

Macroxus similis

(J. E. Gray, 1867)

Family

Sciuridae

Root name

similis

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1843.1.12.54

Type kind

lectotype

Type specimen URI

https://data.nhm.ac.uk/object/3afc71a5-eaff-4524-8f98-cb681ab18e81

Authority page

281

Authority page URI

https://www.biodiversitylibrary.org/page/15624238

Authority publication

Annals and Magazine of Natural History

Name usages

Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)

Sciurus lokriodes

(Bonhote, 1906)

Family

Sciuridae

Root name

lokriodes

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

338

Authority page URI

https://www.biodiversitylibrary.org/page/51223005

Authority publication

Annals and Magazine of Natural History

Name usages

Bonhote (1906:338, https://www.biodiversitylibrary.org/page/51223005) (information at https://hesperomys.com/a/19894)

Sciurus lokriodes Mearsi

(Bonhote, 1906)

Family

Sciuridae

Root name

mearsi

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1906.7.5.10

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/c55c2fca-1b6b-434d-b1c9-4f42901a2ac3

Authority page

338

Authority page URI

https://www.biodiversitylibrary.org/page/51223005

Authority publication

Annals and Magazine of Natural History

Name usages

Bonhote (1906:338, https://www.biodiversitylibrary.org/page/51223005) (information at https://hesperomys.com/a/19894)

Sciurus lokrioides

(O. Thomas, 1908)

Family

Sciuridae

Root name

lokrioides

Validity status

synonym

Nomenclatural status

name_combination · incorrect_subsequent_spelling

Authority page

246

Authority page URI

https://www.biodiversitylibrary.org/page/2222283

Authority publication

Journal of the Bombay Natural History Society

Name usages

Thomas (1908:246, https://www.biodiversitylibrary.org/page/2222283) (information at https://hesperomys.com/a/62341)

Sciurus stevensi

(O. Thomas, 1908)

Family

Sciuridae

Root name

stevensi

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1907.11.26.2

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/de18561c-980b-4dc8-8e43-77692b3c2865

Authority page

246

Authority page URI

https://www.biodiversitylibrary.org/page/2222283

Authority publication

Journal of the Bombay Natural History Society

Name usages

Thomas (1908:246, https://www.biodiversitylibrary.org/page/2222283) (information at https://hesperomys.com/a/62341)

Sciurus pygerythrus janetta

(O. Thomas, 1914)

Family

Sciuridae

Root name

janetta

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1914.12.1.4

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/75ac0cd7-b099-4da8-a259-6a21aea16a1b

Authority page

203

Authority page URI

https://www.biodiversitylibrary.org/page/30453572

Authority publication

Journal of the Bombay Natural History Society

Tomeutes janetta

(O. Thomas, 1915)

Family

Sciuridae

Root name

janetta

Validity status

synonym

Nomenclatural status

name_combination

Authority page

386

Authority page URI

https://www.biodiversitylibrary.org/page/15613112

Authority publication

Annals and Magazine of Natural History

Name usages

Thomas (1915:386, https://www.biodiversitylibrary.org/page/15613112) (information at https://hesperomys.com/a/19890)

Tomeutes lokroides

(O. Thomas, 1915)

Family

Sciuridae

Root name

lokroides

Validity status

synonym

Nomenclatural status

name_combination

Authority page

386

Authority page URI

https://www.biodiversitylibrary.org/page/15613112

Authority publication

Annals and Magazine of Natural History

Name usages

  1. Thomas (1915:386, https://www.biodiversitylibrary.org/page/15613112) (information at https://hesperomys.com/a/19890)
  2. Thomas (1918:371, https://www.biodiversitylibrary.org/page/5454141) (information at https://hesperomys.com/a/32316)
  3. Robinson & Kloss (1918:223, https://www.biodiversitylibrary.org/page/11127045) (information at https://hesperomys.com/a/36143)

Tomeutes pygerythrus

(O. Thomas, 1915)

Family

Sciuridae

Root name

pygerythrus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

386

Authority page URI

https://www.biodiversitylibrary.org/page/15613112

Authority publication

Annals and Magazine of Natural History

Name usages

  1. Thomas (1915:386, https://www.biodiversitylibrary.org/page/15613112) (information at https://hesperomys.com/a/19890)
  2. Robinson & Kloss (1918:226, https://www.biodiversitylibrary.org/page/11127048) (information at https://hesperomys.com/a/36143)

Tomeutes similis

(O. Thomas, 1915)

Family

Sciuridae

Root name

similis

Validity status

synonym

Nomenclatural status

name_combination

Authority page

386

Authority page URI

https://www.biodiversitylibrary.org/page/15613112

Authority publication

Annals and Magazine of Natural History

Name usages

  1. Thomas (1915:386, https://www.biodiversitylibrary.org/page/15613112) (information at https://hesperomys.com/a/19890)
  2. Thomas (1918:371, https://www.biodiversitylibrary.org/page/5454141) (information at https://hesperomys.com/a/32316)

Tomeutes mearsi bellona

(O. Thomas & Wroughton, 1916)

Family

Sciuridae

Root name

bellona

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1915.5.5.177

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/ec45c30c-7951-4885-9c18-59624a72a409

Authority page

420

Authority page URI

https://www.biodiversitylibrary.org/page/30151991

Authority publication

Journal of the Bombay Natural History Society

Name usages

  1. Robinson & Kloss (1918:224, https://www.biodiversitylibrary.org/page/11127046) (information at https://hesperomys.com/a/36143)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)

Tomeutes similis owensi

(O. Thomas & Wroughton, 1916)

Family

Sciuridae

Root name

owensi

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1915.5.5.189

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/96b98e90-7b82-4cf2-b9ae-ef6e78ceb7d1

Authority page

236

Authority page URI

https://www.biodiversitylibrary.org/page/30151787

Authority publication

Journal of the Bombay Natural History Society

Tomeutes mearsi virgo

(O. Thomas & Wroughton, 1916)

Family

Sciuridae

Root name

virgo

Validity status

synonym

Nomenclatural status

available

Type

BMNH:Mamm:1915.5.5.169

Type kind

holotype

Type specimen URI

https://data.nhm.ac.uk/object/dce70967-a327-4540-a044-862fefe3c8ac

Authority page

421

Authority page URI

https://www.biodiversitylibrary.org/page/30151992

Authority publication

Journal of the Bombay Natural History Society

Name usages

  1. Robinson & Kloss (1918:224, https://www.biodiversitylibrary.org/page/11127046) (information at https://hesperomys.com/a/36143)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)

Tomeutes pygerythrus janetta

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

janetta

Validity status

synonym

Nomenclatural status

name_combination

Authority page

226

Authority page URI

https://www.biodiversitylibrary.org/page/11127048

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:226, https://www.biodiversitylibrary.org/page/11127048) (information at https://hesperomys.com/a/36143)

Tomeutes lokroides lokroides

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

lokroides

Validity status

synonym

Nomenclatural status

name_combination

Authority page

223

Authority page URI

https://www.biodiversitylibrary.org/page/11127045

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:223, https://www.biodiversitylibrary.org/page/11127045) (information at https://hesperomys.com/a/36143)

Tomeutes mearsi

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

mearsi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

224

Authority page URI

https://www.biodiversitylibrary.org/page/11127046

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:224, https://www.biodiversitylibrary.org/page/11127046) (information at https://hesperomys.com/a/36143)

Tomeutes mearsi mearsi

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

mearsi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

224

Authority page URI

https://www.biodiversitylibrary.org/page/11127046

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:224, https://www.biodiversitylibrary.org/page/11127046) (information at https://hesperomys.com/a/36143)

Tomeutes lokroides owensi

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

owensi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

224

Authority page URI

https://www.biodiversitylibrary.org/page/11127046

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:224, https://www.biodiversitylibrary.org/page/11127046) (information at https://hesperomys.com/a/36143)

Tomeutes pygerythrus pygerythrus

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

pygerythrus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

226

Authority page URI

https://www.biodiversitylibrary.org/page/11127048

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:226, https://www.biodiversitylibrary.org/page/11127048) (information at https://hesperomys.com/a/36143)

Tomeutes stevensi

(H. C. Robinson & Kloss, 1918)

Family

Sciuridae

Root name

stevensi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

225

Authority page URI

https://www.biodiversitylibrary.org/page/11127047

Authority publication

Records of the Indian Museum

Name usages

Robinson & Kloss (1918:225, https://www.biodiversitylibrary.org/page/11127047) (information at https://hesperomys.com/a/36143)

Tomeutes blythi

(O. Thomas, 1918)

Family

Sciuridae

Root name

blythi

Validity status

synonym

Nomenclatural status

incorrectsubsequentspelling

Authority page

371

Authority page URI

https://www.biodiversitylibrary.org/page/5454141

Authority publication

Journal of the Bombay Natural History Society

Name usages

Thomas (1918:371, https://www.biodiversitylibrary.org/page/5454141) (information at https://hesperomys.com/a/32316)

Callosciurus pygerythrus

J. C. Moore, 1959

Family

Sciuridae

Root name

pygerythrus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

202

Authority publication

Bulletin of the American Museum of Natural History

Name usages

  1. Moore (1959:202) (information at https://hesperomys.com/a/6420)
  2. Moore & Tate (1965:213, https://www.biodiversitylibrary.org/page/2765018) (information at https://hesperomys.com/a/36138)
  3. Corbet & Hill (1980:132) (information at https://hesperomys.com/a/63069)
  4. Honacki, Kinman & Koeppl (1982:348) (information at https://hesperomys.com/a/63071)
  5. Corbet & Hill (1991:140) (information at https://hesperomys.com/a/63070)
  6. Hoffmann, Anderson, Thorington & Heaney (1993:423) (information at https://hesperomys.com/a/69019)
  7. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  8. Smith & Xie (2008:184) (information at https://hesperomys.com/a/64149)
  9. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)
  10. Mammal Diversity Database (2018:ID #3604) (information at https://hesperomys.com/a/67336)
  11. Mammal Diversity Database (2019:ID #3604) (information at https://hesperomys.com/a/67337)
  12. Mammal Diversity Database (2024, https://www.mammaldiversity.org/taxon/1001548) (information at https://hesperomys.com/a/67250)
  13. International Union for the Conservation of Nature (2024, https://www.iucnredlist.org/species/3604/22253451) (information at https://hesperomys.com/a/67246)

Callosciurus pygerythrus blythi

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

blythi

Validity status

synonym

Nomenclatural status

name_combination · incorrect_subsequent_spelling

Authority page

226

Authority page URI

https://www.biodiversitylibrary.org/page/2765031

Authority publication

Fieldiana Zoology

Name usages

Moore & Tate (1965:226, https://www.biodiversitylibrary.org/page/2765031) (information at https://hesperomys.com/a/36138)

Callosciurus pygerythrus janetta

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

janetta

Validity status

synonym

Nomenclatural status

name_combination

Authority page

218

Authority page URI

https://www.biodiversitylibrary.org/page/2765023

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:218, https://www.biodiversitylibrary.org/page/2765023) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  3. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

Callosciurus pygerythrus lokroides

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

lokroides

Validity status

synonym

Nomenclatural status

name_combination

Authority page

227

Authority page URI

https://www.biodiversitylibrary.org/page/2765032

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:227, https://www.biodiversitylibrary.org/page/2765032) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  3. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

Callosciurus pygerythrus mearsi

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

mearsi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

222

Authority page URI

https://www.biodiversitylibrary.org/page/2765027

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:222, https://www.biodiversitylibrary.org/page/2765027) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  3. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

Callosciurus pygerythrus owensi

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

owensi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

220

Authority page URI

https://www.biodiversitylibrary.org/page/2765025

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:220, https://www.biodiversitylibrary.org/page/2765025) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  3. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

Callosciurus pygerythrus pygerythrus

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

pygerythrus

Validity status

synonym

Nomenclatural status

name_combination

Authority page

217

Authority page URI

https://www.biodiversitylibrary.org/page/2765022

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:217, https://www.biodiversitylibrary.org/page/2765022) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)

Callosciurus pygerythrus stevensi

J. C. Moore & Tate, 1965

Family

Sciuridae

Root name

stevensi

Validity status

synonym

Nomenclatural status

name_combination

Authority page

224

Authority page URI

https://www.biodiversitylibrary.org/page/2765029

Authority publication

Fieldiana Zoology

Name usages

  1. Moore & Tate (1965:224, https://www.biodiversitylibrary.org/page/2765029) (information at https://hesperomys.com/a/36138)
  2. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  3. Smith & Xie (2008:184) (information at https://hesperomys.com/a/64149)
  4. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

Callosciurus pygerythrus blythii

Thorington & R. S. Hoffmann, 2005

Family

Sciuridae

Root name

blythii

Validity status

synonym

Nomenclatural status

name_combination

Authority publication

Baltimore

Name usages

  1. Thorington & Hoffmann (2005) (information at https://hesperomys.com/a/8554)
  2. Wilson, Lacher & Mittermeier (2016:733) (information at https://hesperomys.com/a/59599)

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